Antibacterial substance found in lichens. Isoln from varieties of Usnea barbata (L.) Wigg., Usneaceae: Rochleder, Heldt, Ann. 48, 11 (1843); Widman, Ann. 310, 230 (1900); 324, 139 (1902). Isoln from Ramalina reticulata: Marshak, Public Health Rep. 62, 3 (1947); Stark et al., J. Am. Chem. Soc. 72, 1819 (1950). Occurs in nature in both the d- and l-forms as well as a racemic mixture. Structure: Curd, Robertson, J. Chem. Soc. 1937, 894; Schpf, Ross, Ann. 546, (1941); Barton, Brunn, J. Chem. Soc. 1953, 603. Resolution of (±)-usnic acid: Dean et al., ibid. 1250. Synthesis: Barton et al., ibid. 1956, 530. Biosynthesis in vitro: Penttila, Fales, Chem. Commun. 1966, 656. Absolute configuration of (+)-form: S. Huneck et al., Tetrahedron Lett. 22, 351 (1981). Toxicity study: Antibiotics I, D. Gottlieb, P. Shaw, Eds. (Springer Verlag, New York, 1967) p 611.